There are other ideas I have in mind but I tried to draw out at least one of the 5 kriggle subspecies's male form.
Unlike the females, the males can't fly very fast; instead they float and can stay in the air much longer than the females (imagine a kite vs an air balloon) Males also have their strength more in their legs than their arms, allowing them to carry heavy cargo across normally trek-less lands. Also, their neck sports an inflation organ to allow such buoyancy.
Let me know what you guys think, and feel free to add input.